References

as of 09/2022 with “Cardiocondyla” in the title or abstract, starting with August Forel’s description of the wingless males and intranidal mating in 1892, or as he termed it ‘reproduction consanguine perpétuée’

124. Jaimes-Niño LM, Süß A, Heinze J, Schultner E, Oettler J (preprint) The indispensable soma of Cardiocondyla obscurior ants. bioRxiv 2022.10.02.510526. https://doi.org/10.1101/2022.10.02.510526

123. Schultner E, Wallner T, Dofka B, Brülhart J, Heinze J, Freitak D, Pokorny T, Oettler J. Queens control caste allocation in the ant Cardiocondyla obscurior. Proceedings of the Royal Society B: Biological Sciences. 2023;290:20221784. https://doi.org/10.1098/rspb.2022.1784

122. Vidal M, Heinze J. Assisted dispersal and reproductive success in an ant species with matchmaking. Ecology and Evolution. 2022;12(8):e9236. https://doi.org/10.1002/ece3.9236

121. Jaimes-Niño LM, Heinze J, Oettler J. Late-life fitness gains and reproductive death in Cardiocondyla obscurior ants. eLife. 2022;11:e74695. https://doi.org/10.7554/eLife.74695

120. Jackson R, Monnin D, Patapiou PA, Golding G, Helanterä H, Oettler J, Heinze J, Wurm Y, Economou CK, Chapuisat M, Henry LM. Convergent evolution of a labile nutritional symbiosis in ants. The ISME Journal. 2022;16:2114-2122. https://doi.org/10.1038/s41396-022-01256-1.

119. Leponiemi M, Schultner E, Dickel F, Freitak D. Chronic sublethal pesticide exposure affects brood production, morphology, and endosymbionts, but not immunity in the ant, Cardiocondyla obscurior. Ecological Entomology. 2022;47(3):273-283. https://doi.org/10.1111/een.13111.

118. Ün Ç, Hacker J, Heinze J, Oettler J, Schultner E. Endosymbionts mediate the effects of antibiotic exposure in the tramp ant Cardiocondyla obscurior. Ecological Entomology. 2022;47(2):170-180. https://doi.org/10.1111/een.13099.

117. Majeed W, Koch EBA, Rana N, Naseem R. First Record of Cardiocondyla obscurior Wheeler, 1929 (Hymenoptera: Formicidae: Myrmicinae) for Pakistan. Sociobiology. 2021;68(3):e5907.

116. Errbii M, Keilwagen J, Hoff KJ, Steffen R, Altmüller J, Oettler J, Schrader L. Transposable elements and introgression introduce genetic variation in the invasive ant Cardiocondyla obscurior. Molecular Ecology. 2021;30(23):6211-6228. https://doi.org/10.1111/mec.16099

115. Heinze J, Marschall J, Lautenschläger B, Seifert B, Gratiashvili N, Strohm E. Courtship with two spoons-Anatomy and presumed function of the bizarre antennae of Cardiocondyla zoserka ant males. Ecology and Evolution. 2021;11(12):7827–7833.

114. Harrison MC, Niño LMJ, Rodrigues MA, Ryll J, Flatt T, Oettler J, Bornberg-Bauer E. Gene coexpression network reveals highly conserved, well-regulated anti-ageing mechanisms in old ant queens. Genome Biology and Evolution. 2021;13(6):evab093. https://doi.org/10.1093/gbe/evab093

113. Ün Ç, Schultner E, Manzano-Marín A, Flórez LV, Seifert B, Heinze J, Oettler J. Cytoplasmic incompatibility between Old and New World populations of a tramp ant. Evolution. 2021;75:1775–1791.  https://doi.org/10.1111/evo.14261

112. Vidal M, Königseder F, Giehr J, Schrempf A, Lucas C, Heinze J. Worker ants promote outbreeding by transporting young queens to alien nests. Communications Biology. 2021;4(1):515.

111. Heinze J, Giehr J. The plasticity of lifespan in social insects. Philosophical Transactions of the Royal Society B: Biological Sciences. 2021;376(1823):20190734.

110. Schrader L, Winter M, Errbii M, Delabie J, Oettler J, Gadau J. Inhibition of HSP90 causes morphological variation in the invasive ant Cardiocondyla obscurior. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution. 2021;336:333–340.

109. Oettler J. Cardiocondyla: Heart Node Ants. In: Starr C, editor. Encyclopedia of Social Insects. 2020. pp. 1–4.

108. Heinze J, Trindl A, Seifert B, Ma MK, Maung W. First records of Cardiocondyla ants from Myanmar. Asian Myrmecology. 2020;12:1–8.

107. Heinze J. Emeryia, Xenometra, zoserka: it’s a boy, again! The misleading morphology of Cardiocondyla male ants. Insect Sociaux. 2019;67(1):139–146.

106. Jacobs S, Heinze J. Population and colony structure of an ant with territorial males, Cardiocondyla venustula. BMC Evolutionary Biology. 2019;19(1):115–119.

105. Liu L, Wu Y, Chen F, Wang Q-X, Zhang X-Y, Tang Y, Li F, Qian Z-Q. Characterization of the complete mitochondrial genome of the invasive tramp ant Cardiocondyla obscurior (Hymenoptera: Formicidae: Myrmicinae). Mitochondrial DNA Part B. 2019;4(1):1496–1498.

104. Heinze J, Frohschammer S, Bernadou A. When invasive ants meet: effects of outbreeding on queen performance in the tramp ant Cardiocondyla itsukii. Insect Science. 2019;26(2):333–340.

103. Oettler J, Platschek T, Schmidt C, Rajakumar R, Favé M-J, Khila A, Abouheif E. Interruption points in the wing gene regulatory network underlying wing polyphenism evolved independently in male and female morphs in Cardiocondyla ants. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution. 2019;332(1-2):7–16.

102. Heinze J, Hanoeffner M, Delabie JHC, Schrempf A. Methuselah’s daughters: Paternal age has little effect on offspring number and quality in Cardiocondyla ants. Ecology and Evolution. 2018;8(23):12066–12072.

101. Christine S V, Jürgen H. Mandible morphology reflects the type of male antagonism in the ant genus Cardiocondyla (Hymenoptera: Formicidae). European Journal of Entomology; 2018;115:455–466.

100. Fuessl M, Santos CG, Hartfelder K, Schrempf A, Heinze J. Accessory gland proteins of males in the male-diphenic ant Cardiocondyla obscurior. Physiological Entomology. 2018;43(4):276–284.

99. Metzler S, Schrempf A, Heinze J. Individual- and ejaculate-specific sperm traits in ant males. Journal of Insect Physiology 2018;107:284–290.

98. Giehr J, Heinze J, Schrempf A. Group demography affects ant colony performance and individual speed of queen and worker aging. BMC Evolutionary Biology. 2017;17(1):173.

97. Seifert B, Okita I, Heinze J. A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae). Zootaxa. 2017;4290(2):324–356.

96. Giehr J, Grasse AV, Cremer S, Heinze J, Schrempf A. Ant queens increase their reproductive efforts after pathogen infection. Royal Society Open Science 2017;4(7):170547.

95. Schmidt CV, Heinze J. Genital morphology of winged and wingless males in the ant genus Cardiocondyla (Formicidae, Myrmicinae). Insect Systematics & Evolution. 2017;49(1):59–80.

94. Heinze J. Life-history evolution in ants: the case of Cardiocondyla. Proceedings of the Royal Society B: Biological Sciences. 2017;284:20161406.

93. Schrader L, Helanterä H, Oettler J. Accelerated evolution of developmentally biased genes in the tetraphenic ant Cardiocondyla obscurior. Molecular Biology and Evolution. 2017;34(3):535–544.

92. Schmidt CV, Heimbucher A, Bernadou A, Heinze J. First come, first served: the first-emerging queen monopolizes reproduction in the ant Cardiocondyla “argyrotricha.” Journal of Ethology. 2017 ;35(1):21–27.

91. Schrempf A, Giehr J, Röhrl R. Royal darwinian demons: Enforced changes in reproductive efforts do not affect the life expectancy of ant queens. The American Naturalist. 2017;189:436–442.

90. Seifert B, Okita I, Heinze J. A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae). Zootaxa. 2017;4290:324–356.

89. Metzler S, Heinze J, Schrempf A. Mating and longevity in ant males. Ecology and Evolution. 2016;6(24):8903–8906.

88. Schrempf A, Moser A, Delabie J, Heinze J. Sperm traits differ between winged and wingless males of the ant Cardiocondyla obscurior. Integrative Zoology. 2016;11(6):427–432.

87. Oettler J, Schrempf A. Fitness and aging in Cardiocondyla obscurior ant queens. Current Opinion in Insect Science. 2016;16:58–63.

86. Heinze J, Seifert B, Zieschank V. Massacre of the innocents in Cardiocondyla thoracica: Manipulation by adult ant males incites workers to kill their immature rivals. Entomological Science. 2016;19(3):239–244.

85. Heinze J. The male has done his work–the male may go. Current Opinion in Insect Science. 2016;16:22–26.

84. Jacobs S, Heinze J. Between fighting and tolerance: reproductive biology of wingless males in the ant Cardiocondyla venustula. Insect Science. 2017;24(5):818-828.

83. Wyschetzki von K, Lowack H, Heinze J. Transcriptomic response to injury sheds light on the physiological costs of reproduction in ant queens. Molecular Ecology. 2016;25(9):1972–1985.

82. Okita I, Tsuchida K. Clonal reproduction with androgenesis and somatic recombination: the case of the ant Cardiocondyla kagutsuchi. Naturwissenschaften. 2016;103(3-4):22.

81. Klein A, Schultner E, Lowak H, Schrader L, Heinze J, Holman L, Oettler J. Evolution of social insect polyphenism facilitated by the sex differentiation cascade. PLoS Genetics. 2016;12(3):e1005952.

80. Schmidt CV, Schrempf A, Trindl A, Heinze J. Microsatellite markers for the tramp ant, Cardiocondyla obscurior (Formicidae: Myrmicinae). Journal of Genetics. 2016;95(1):e1–4.

79. Wyschetzki von K, Rueppell O, Oettler J, Heinze J. Transcriptomic signatures mirror the lack of the cecundity/longevity trade-off in ant queens. Molecular Biology and Evolution. 2015;32(12):3173–3185.

78. Rueppell O, Königseder F, Heinze J, Schrempf A. Intrinsic survival advantage of social insect queens depends on reproductive activation. Journal of Evolutionary Biology. 2015;28(12):2349–2354.

77. Okita I, Terayama M, Tsuchida K. Cryptic Lineages in the Cardiocondyla sl. kagutsuchi Terayama (Hymenoptera: Formicidae) discovered by phylogenetic and morphological approaches. Sociobiology. 2015;62(3):401.

76. Schrempf A, Wyschetzki von K, Klein A, Schrader L, Oettler J, Heinze J. Mating with an allopatric male triggers immune response and decreases longevity of ant queens. Molecular Ecology. 2015;24(14):3618–3627.

75. Schrader L, Simola DF, Heinze J, Oettler J. Sphingolipids, transcription factors, and conserved toolkit genes: Developmental plasticity in the ant Cardiocondylaobscurior. Molecular Biology and Evolution. 2015;32(6):1474–1486.

74. Klein A, Schrader L, Gil R, Manzano-Marín A, Flórez L, Wheeler D, Werren JH, Latorre A, Heinze J, Kaltenpoth M, Moya A, Oettler J. A novel intracellular mutualistic bacterium in the invasive ant Cardiocondyla obscurior. The ISME Journal. 2015;10(2):376–388.

73. Oettler J, Nachtigal A-L, Schrader L. Expression of the foraging gene is associated with age polyethism, not task preference, in the ant Cardiocondyla obscurior. PLoS ONE. 2015;10(12):e0144699.

72. Kramer BH, Schrempf A, Scheuerlein A, Heinze J. Ant Colonies Do Not Trade-Off Reproduction against Maintenance. PLoS ONE. 2015;10(9):e0137969.

71. Fuessl M, Schrempf A, Heinze J. Queen and male longevity in the Southeast Asian ant Cardiocondyla tjibodana Karavaiev, 1935. Asian Myrmecology. 2015;7:137–141.

70. Fuessl M, Reinders J, Oefner PJ, Heinze J, Schrempf A. Selenophosphate synthetase in the male accessory glands of an insect without selenoproteins. Journal of Insect Physiology. 2014;71:46–51.

69. Heinze J, Schrempf A, Wanke T, Rakotondrazafy H, Rakotondrazafy T, Fisher BL. Polygyny, Inbreeding and Wingless Males in the Malagasy Ant Cardiocondyla shuckardi Forel (Hymenoptera, Formicidae). Sociobiology. 2014;6:300–306.

68. Schrempf A. Inbreeding, multiple mating and foreign sexuals in the ant Cardiocondyla nigra (Hymenoptera: Formicidae). Myrmecological News. 2014;20:1–5.

67. Schrader L, Kim JW, Ence D, Zimin A, Klein A, Wyschetzki K, Weichselgartner T, Kemene C, Stöckl J, Schultner E, Wurm Y, Smith CD, Yandell M, Heinze J, Gadau J, Oettler J. Transposable element islands facilitate adaptation to novel environments in an invasive species. Nature Communications. 2014;5:5495.

66. Bressan JMA, Benz M, Oettler J, Heinze J, Hartenstein V, Sprecher SG. A map of brain neuropils and fiber systems in the ant Cardiocondyla obscurior. Frontiers in Neuroanatomy. 2014;8:166.

65. Schmidt CV, Frohschammer S, Schrempf A, Heinze J. Virgin ant queens mate with their own sons to avoid failure at colony foundation. Naturwissenschaften. 2013;101(1):69–72.

64. Heinze J, Frohschammer S, Bernadou A. Queen life-span and total reproductive success are positively associated in the ant Cardiocondyla cf. kagutsuchi. Behavioral Ecology and Sociobiology. 2013;67(10):1555–1562.

63. Seifert B, Frohschammer S. Cardiocondyla pirata sp. n. – a new Philippine ant with enigmatic pigmentation pattern (Hymenoptera, Formicidae). Zookeys. 2013;(301):13–24.

62. Okita I, Terayama M. The spatial distribution of mtDNA and phylogeographic analysis of the ant Cardiocondyla kagutsuchi (Formicidae) in Japan. Sociobiology. 2013;60:129–134.

61. Will S, Delabie JHC, Heinze J, Ruther J, Oettler J. Cuticular lipid profiles of fertile and non-fertile Cardiocondyla ant queens. Journal of Insect Physiology. 2012;58(9):1245–1249.

60. Wetterer JK, Wetterer. Worldwide spread of Emery’s sneaking ant, Cardiocondyla emeryi (Hymenoptera: Formicidae). Myrmecological News. 2012;17:13-20.

59. Heinze J, Aumeier V, Bodenstein B, Crewe RM, Schrempf A. Wingless and intermorphic males in the ant Cardiocondyla venustula. Insect Sociaux. 2012;60(1):43–48.

58. Wetterer JK. Worldwide spread of the moorish sneaking ant, Cardiocondyla mauritanica (Hymenoptera: Formicidae). Sociobiology. 2012;59(3):985–997.

57. Heinze J, Schrempf A. Terminal investment: individual reproduction of ant queens increases with age. PLoS ONE. 2012;7(4):e35201.

56. Cremer S, Suefuji M, Schrempf A, Heinze J. The dynamics of male-male competition in Cardiocondyla obscurior ants. BMC Ecology. 2011;12:7–7.

55. Yoshizawa J, Yamauchi K, Tsuchida K. Decision-making conditions for intra- or inter-nest mating of winged males in the male-dimorphic ant Cardiocondyla minutior. Insect Sociaux. 2011;58(4):531–538.

54. Schrempf A, Cremer S, Heinze J. Social influence on age and reproduction: reduced lifespan and fecundity in multi-queen ant colonies. Journal of Evolutionary Biology. 2011;24(7):1455–1461.

53. Cremer S, Schrempf A, Heinze J. Competition and opportunity shape the reproductive tactics of males in the ant Cardiocondyla obscurior. PLoS ONE. 2011;6(3):e17323.

52. Heinze J, Weber M. Lethal sibling rivalry for nest inheritance among virgin ant queens. Journal of Ethology. 2011;29:197–201.

51. Oettler J, Suefuji M, Heinze J. The evolution of alternative reproductive tactics in male Cardiocondyla ants. Evolution. 2010;64(11):3310–3317.

50. Ugelvig LV, Kronauer DJC, Schrempf A, Heinze J, Cremer S. Rapid anti-pathogen response in ant societies relies on high genetic diversity. Proceedings of the Royal Society B: Biological Sciences. 2010;277(1695):2821–2828.

49. Heinze J, Christine S V, Nugroho H, Seifert B. Wingles fighter males in the Wallacean ant Cardiocondyla nigrocerea (Insecta: Formicidae). The Raffles Bulletin of Zoology. 2010;58:1–6.

48. Frohschammer S, Heinze J. A heritable component in sex ratio and caste determination in a Cardiocondyla ant. Frontiers in Zoology. 2009;6:27.

47. Suefuji M, Cremer S, Oettler J, Heinze J. Queen number influences the timing of the sexual production in colonies of Cardiocondyla ants. Biology Letters. 2008;4(6):670–673.

46. Frohschammer S, Heinze J. Male fighting and “territoriality” within colonies of the ant Cardiocondyla venustula. Naturwissenschaften. 2008;96(1):159–163.

45. Yoshizawa J, Mimori K, Yamauchi K, Tsuchida K. Sex mosaics in a male dimorphic ant Cardiocondyla kagutsuchi. Naturwissenschaften. 2008;96(1):49–55.

44. Cremer S, D’Ettorre P, Drijfhout FP, Sledge MF, Turillazzi S, Heinze J. Imperfect chemical female mimicry in males of the ant Cardiocondyla obscurior. Naturwissenschaften. 2008;95(11):1101–1105.

43. Schrempf A, Heinze J. Mating with stressed males increases the fitness of ant queens. PLoS ONE. 2008;3(7):e2592.

42. Seifert B. Cardiocondyla atalanta Forel, 1915, a cryptic sister species of Cardiocondyla nuda (Mayr, 1866) (Hymenoptera: Formicidae). Myrmecological News. 2008;11:43–48.

41. Mercier JL, Lenoir JC, Eberhardt A, Frohschammer S, Williams C, Heinze J. Hammering, mauling, and kissing: stereotyped courtship behavior in Cardiocondyla ants. Insect Sociaux. 2007;54(4):403–411.

40. Yamauchi K, Ishida Y, Hashim R, Heinze J. Queen-queen competition and reproductive skew in a Cardiocondyla ant. Insect Sociaux. 2007;54(3):268–274.

39. Schrempf A, Heinze J. Back to one: consequences of derived monogyny in an ant with polygynous ancestors. Journal of Evolutionary Biology 2007;20(2):792–799.

38. Schrempf A, Darrouzet E, Heinze J. Mating success and potential male-worker conflict in a male-dimorphic ant. BMC Evolutionary Biology. 2007;7(1):114.

37. Du Y, Schrempf A, Heinze J. Environmental determination of the male morph in the ant Cardiocondyla obscurior (Hymenoptera : Formicidae). European Journal of Entomology. 2007;104(2):243–246.

36. Yamauchi K, Ishida Y, Hashim R, Heinze J. Queen-queen competition by precocious male production in multiqueen ant colonies. Current Biology. 2006;16(24):2424–2427.

35. Lenoir JC, Schrempf A, Lenoir A, Heinze J, Mercier JL. Genetic structure and reproductive strategy of the ant Cardiocondyla elegans: strictly monogynous nests invaded by unrelated sexuals. Molecular Ecology. 2006;16(2):345–354.

34. Schrempf A, Aron S, Heinze J. Sex determination and inbreeding depression in an ant with regular sib-mating. Heredity. 2006;97(1):75–80.

33. Schrempf A, Heinze J. Proximate mechanisms of male morph determination in the ant Cardiocondyla obscurior. Evolution and Development. 2006;8(3):266–272.

32. Heinze J, Cremer S, Eckl N, Schrempf A. Stealthy invaders: the biology of Cardiocondyla tramp ants. Insect Sociaux. 2006;53:1–7.

31. Heinze J, Delabie J. Population structure of the male-polymorphic ant Cardiocondyla obscurior. Studies on Neotropical Fauna and Environment. 2005;40(3):187–190.

30. Heinze J, Trindl A, Seifert B, Yamauchi K. Evolution of male morphology in the ant genus Cardiocondyla. Molecular Phylogenetics and Evolution. 2005;37(1):278–288.

29. Yamauchi K, Asano Y, Lautenschläger B, Trindl A, Heinze J. A new type of male dimorphism with ergatoid and short-winged males in Cardiocondyla cf. kagutsuchi. Insect Sociaux. 2005;52(3):274–281.

28. Schrempf A, Heinze J, Cremer S. Sexual cooperation: mating increases longevity in ant queens. Current Biology. 2005;15(3):267–270.

27. de Menten L, Cremer S, Heinze J, Aron S. Primary sex ratio adjustment by ant queens in response to local mate competition. Animal Behaviour. 2005;69:1031–1035.

26. Schrempf A, Reber C, Tinaut A, Heinze J. Inbreeding and local mate competition in the ant Cardiocondyla batesii. Behavioral Ecology and Sociobiology. 2004;57(5):502–510.

25. Heinze J, Böttcher A, Cremer S. Production of winged and wingless males in the ant, Cardiocondyla minutior. Insect Sociaux. 2004;51(3).

24. Seifert B. The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae)-a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi and C. minutior species groups. Annalen des Naturhistorischen Museums in Wien. Serie B für Botanik und Zoologie. 2003;104(B):203–338.

23. Cremer S, Heinze J. Stress grows wings: Environmental induction of winged dispersal males in Cardiocondyla ants. Current Biology. 2003;13(3):219–223.

22. Anderson C, Cremer S, Heinze J. Live and let die: why fighter males of the ant Cardiocondyla kill each other but tolerate their winged rivals. Behavioral Ecology. 2003;14(1):54–62.

21. Cremer S, Sledge MF, Heinze J. Chemical mimicry: male ants disguised by the queen’s bouquet. Nature. 2002;419(6910):897.

20. Heinze J, Schrempf A, Seifert B, Tinaut A. Queen morphology and dispersal tactics in the ant, Cardiocondyla batesii. Insect Sociaux. 2002;49(2):129–132.

19. Cremer S, Heinze J. Adaptive production of fighter males: queens of the ant Cardiocondyla adjust the sex ratio under local mate competition. Proceedings of the Royal Society B: Biological Sciences. 2002;269(1489):417–422.

18. Cremer S, Lautenschläger B, Heinze J. A transitional stage between the ergatoid and winged male morph in the ant Cardiocondyla obscurior. Insect Sociaux. 2002;49(3):221–228.

17. Heinze J, Hölldobler B, Yamauchi K. Male competition in Cardiocondyla ants. Behavioral Ecology and Sociobiology. 1998;42(4):239–246.

16. Collingwood C, Tigar B, Agosti D. Introduced ants in the United Arab Emirates. Journal of Arid Environments. 1997;37(3):505–512.

15. Heinze J, Trenkle S. Male polymorphism and gynandromorphs in the ant Cardiocondyla emeryi. Naturwissenschaften. 1997;84(3):129–131.

14. MacKay W. New distributional records for the ant genus Cardiocondyla in the new world (Hymenoptera: Formicidae. Pan Pacific Entomologist. 1995;71:1–4.

13. Tsuji N, Yamauchi K, Yamamura N. A mathematical model for wing dimorphism in male Cardiocondyla ants. Journal of Ethology. 1994;12(1):19–24.

12. Heinze J, Hölldobler B. Fighting for a harem of queens: physiology of reproduction in Cardiocondyla male ants. Proceedings of the National Academy of Sciences USA. 1993;90(18):8412–8414.

11. Heinze J, Kühnholz S, Schilder K, Hölldobler B. Behavior of ergatoid males in the ant, Cardiocondyla nuda. Insect Sociaux. 1993;40(3):273–282.

10. Yamauchi K, Kawase N. Pheromonal manipulation of workers by a fighting male to kill his rival males in the ant Cardiocondyla wroughtonii. Naturwissenschaften. 1992;79(6):274–276.

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